Dr. Shiel received a Bachelor of Science degree with honors from the University of Notre Dame. There he was involved in research in radiation biology and received the Huisking Scholarship. After graduating from St. Louis University School of Medicine, he completed his Internal Medicine residency and Rheumatology fellowship at the University of California, Irvine. He is board-certified in Internal Medicine and Rheumatology.
Alanine: An amino acid, one of the 20 building blocks of protein. It is not essential to the diet, as it can be made by the body from other substances. Alanine was discovered in protein in 1875. Symbol: Ala.