Dr. Shiel received a Bachelor of Science degree with honors from the University of Notre Dame. There he was involved in research in radiation biology and received the Huisking Scholarship. After graduating from St. Louis University School of Medicine, he completed his Internal Medicine residency and Rheumatology fellowship at the University of California, Irvine. He is board-certified in Internal Medicine and Rheumatology.
Cytogenetics: The study of chromosomes, which are the visible carriers of the hereditary material. Cytogenetics is a fusion science, joining cytology (the study of cells) with genetics (the study of inherited variation).